For four years, the biggestStar Trekalien villain crossover has been gradually happening onStar Trek: Lower Decks, but in the animated comedy’s opening credits. Created by Mike McMahan,Star Trek: Lower Decksis set in the late 24th century ofStar Trek: The Next Generation’s era. This means everythingStar Trekup to that point, fromStar Trek: Enterprise, toStar Trek: The Original Series, TNG, Star Trek: Deep Space Nine, Star Trek: Voyager,and the first 10Star Trekmoviesare ripe for homage or parody. And this includes thenumerous aliens Starfleet and the United Federation of Planets have faced.

Star Trek: Lower Decks' opening credits evolve throughout the show’s five seasons. Set to the majestic score by Chris Westlake,Star Trek: Lower Decks' titles showcase the USS Cerritos and its misadventures warping around the galaxy. After the names of the four mainStar Trek: Lower Deckscast members, Tawny Newsome, Jack Quaid, Noël Wells, and Eugene Cordero, the Cerritos runs into a region of space where Starfleet’s enemy aliens are fighting each other. As a California Class support ship,the Cerritos wisely turns tail and runs away.With each season ofStar Trek: Lower Decks, more and more aliens show up for the big fight in the opening credits. Here are all 10 aliens involved inStar Trek: Lower Decks' main titles showdown.

Scott Bakula as Captain Archer, Kate Mulgrew as Captain Janeway, and Patrick Stewart as Captain Picard

5Star Trek: Lower Decks Season 1

The Borg and the Romulans

The Borg.Logically, the USS Cerritos hits the bricks inStar Trek: Lower Decks' opening credits at the sight of several Borg Cubes fighting Romulan Warbirds. The Borg Collective are the most fearsome threat to the United Federation of Planets - and the entire galaxy - in the late 24th century.InStar Trek: Lower Decks' timeframe, the Borg are at their mightiest.Led by the malevolentBorg Queen (Alice Krige, Susanna Thompson), the Borg have attacked the Federation multiple times, although they have always been defeated, primarily by Captain Jean-Luc Picard (Patrick Stewart) and the crew of the USS Enterprise-D and E.

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The Romulans.Battling it out with the Borg are several Warbirds belonging to the Romulan Star Empire.One of the Federation’s oldest foes, the Romulans are a secretive and insidious military society. InStar Trek: Lower Decks' time frame, the Romulans have just become aware their sun will go supernova. However, they’re steadfast enemies of the Federation duringLower Decks. Interestingly,Star Trek: Lower Deckspremiered on Paramount+ in 2020 afterStar Trek: Picard, and the sight of Romulans fighting the Borg can be seen as a nod to how the Romulans acquired a derelictBorg Cube and turned it into the Artifact, a Borg Reclamation site, inStar Trek: Picardseason 1.

Star Trek Lower Decks season 3 villains intro

4Star Trek: Lower Decks Season 2

Pakleds and the Klingons

The Pakleds.InStar Trek: Lower Decksseason 2’s opening credits, Pakleds Clump ships and Klingon Birds-of-Prey join the big fight between the Borg and the Romulans. ThePakleds may be the dumbest of the alien foesintroduced inStar Trek: The Next Generation. The Pakleds returned inStar Trek: Lower Decks, and they were still imbeciles, but they were also dangerous.The Pakleds nearly destroyed the USS Cerritos, and they framed Captain Carol Freeman(Dawnn Lewis) for blowing up Pakled Planet.

The Klingon Empire are allies of the Federation.

The Klingons.InStar Trek: Lower Decks' late 24th century,theKlingon Empire are allies of the Federation, and they fought and wonStar Trek: Deep Space Nine’s Dominion War together. TheKlingons are led by Chancellor Gowron(J.G. Hertzler), who became leader of the Empire at the end ofDS9.Klingons and Starfleet may be on friendly political terms, but the warrior race generally frowns upon humans, and many Starfleet Officers don’t enjoy being around Klingons either. But if it’s a good day to die, Klingons will do so besides Starfleet if honor demands it.

3Star Trek: Lower Decks Season 3

The Crystalline Entity

The Crystalline Entity.InStar Trek: Lower Decksseason 3’s opening credits, the overwhelming power of the Borg meets its match in the Crystalline Entity, which joins the fray with the Borg, Romulans, Pakleds, and Klingons. Introduced inStar Trek: The Next Generationseason 1, theCrystalline Entity is a massive, snowflake-shaped organism with the power to wipe out all life on a planet. The first Crystalline Entity was destroyed by the USS Enterprise-D after it was responsible for thousands of deaths. However,Star Trek: Lower Decksseason 2, episode 6, “I, Excretus,” revealed that there are multiple Crystalline Entities in the galaxy.

The Crystalline Entity remains a popular alien life form with TNG fans.

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The original Crystaline Entity first appeared inStar Trek: The Next Generationseason 1’s “Datalore,” when a reactivatedLore (Brent Spiner)tried to feed the crew of the USS Enterprise-D to the Entity. The Crystalline Entity fled after this initial encounter with the Enterprise, but it returned inStar Trek: The Next Generationseason 5’s “Silicon Avatar.” After wiping out a Federation colony on Melona IV, the Enterprise-D attemped to communicate with the Crystalline Entity but it was destroyed by the resonance from graviton emissions. Still,the Crystalline Entity remains a popular alien life form withTNGfans.

2Star Trek: Lower Decks Season 4

The Whale Probe and the Breen

The Whale Probe.InStar Trek: Lower Decksseason 4’s opening credits, the Whale Probe and the Breen join the growing alien conflagration the USS Cerritos flees from. Introduced inStar Trek IV: The Voyage Home,theWhale Probe is an unexplained extraterrestrial entity that nearly destroyed Earth as it searched for humpback whales. Admiral James T. Kirk (William Shatner) and the former crew of the USS Enterprise time-traveled to 1986 San Francisco and brought two humpback whales back to the future. After communicating with the whales, the Probe was satisfied and retreated.

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The Breen.First seen inStar Trek: Deep Space Nine(although previously mentioned inStar Trek: The Next Generation), the Breen Conferacy are mysterious aliens never seen without their helmets and refigeration suits. The Breen allied with the Dominion and nearly conquered the Federation during the Dominion War.The Breen made a cameo inStar Trek: Lower Decks,butStar Trek: Discoveryrevealed more about the renamed Breen Imperium in the 32nd century by introducing L’ak (Elias Toufexis), the Breen Scion and the first of theBreen to be seen without his helmet.

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1Star Trek: Lower Decks Season 5

Tholians, Apollo’s Hand, and V’Ger

The Tholians. InStar Trek: Lower Decks' fifth and final seasonon Paramount+, the opening credits' alien fight turns into an all-out brawl as three more villains join the conflagration: the Tholians, the hand of Apollo (Michael Forest), and V’Ger.The Tholians are an insect-like crystalline racewho first appeared inStar Trek: The Original Series'“The Tholian Web.” Tholians are known for their tactic of creating an energy net spun by their starships. First Contact between Tholians and United Earth occurred inStar Trek: Enterprise, but Tholians are often mentioned yet rarely seen inStar Trek.

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Apollo’s Hand.The Borg Cube inStar Trek: Lower Decks' opening credits is grasped by the unforgettable green hand of Apollo. Apollo used his giant green hand to trap the Starship Enterprise inStar Trek: The Original Seriesseason 2’s “Who Mourns for Adonais,” which revealedan alien being claiming to be the Greek demigod Apollo living on the planet Pollux IV. According to Apollo, he and the Greek pantheon where explorers who visited Earth 5000 years earlier. Although Apollo later claimed he was born on Earth to Zeus and the mortal woman Leto. After Captain Kirk and the Enterprise destroyed Apollo’s power source, the so-called Greek god turned himself into energy and vanished.

Star Trek: Lower Decks Season 5 Official Poster

V’Ger journeyed home to Earth seeking its creator.

V’Ger.Last but not least, V’Ger makes a startling appearance as the final alien villain inStar Trek: Lower Decksseason 5’s opening credits. Introduced inStar Trek: The Motion Picture, V’Ger appeared to be a vast energy cloud of enormous destructive power heading toward Earth. Intercepted by the USS Enterprise commanded by Admiral Kirk, V’Ger was revealed to be a massive sentient starship that was originally the Earth space probe Voyager 6. Rebuilt by living machines, V’Ger journeyed home to Earth seeking its creator. V’Ger combined with Commander Will Decker (Stephen Collins) andLt. Ilia (Persis Khambatta)and ascended into another state of being. But inStar Trek: Lower Decksseason 5’s opening credits, V’Ger is back and tips the scales in the big alien brawl the USS Cerritos warps away from.